Modulatricidae – Spot-throat & Allies
Modulatricidae is a family of just three birds in three different genera.
They were formerly included in Promeropidae until recent DNA studies confirmed their status as related, but a separate family. Previously, some authorities included them in the Muscicapidae, Turdidae or Passeroidea. Nowadays, they are thought to be sister to the sugarbirds, as both are basal Passeroidea.
The Dapple-throat or Dappled Mountain-robin Arcanator orostruthus is found in Mozambique and Tanzania.
The Spot-throat Modulatrix stictigula is found in northern Malawi and Tanzania.

Spot-throat Modulatrix stictigula – ©Dubi Shapiro
The Grey-chested Babbler Kakamega poliothorax, also known as the Grey-chested Illadopsis, is found in Burundi, Cameroon, Democratic Republic of the Congo, Equatorial Guinea, Kenya, Nigeria, Rwanda, and Uganda.
For all three species, their natural habitat is subtropical or tropical moist montane forests. All three are threatened by habitat loss, although the later, being widespread, is least vulnerable. The other two species favour small. Isolated areas, meaning we know little about them and they are more threatened. The nest and eggs of only one of the three species has been described.
They all forage for insects on the ground, in dense undergrowth, and are more often heard than seen. In the case of the Dapple-throat, it favours leaf litter near streams, searching for invertebrates. Their whistled songs and calls has been compared to those of Old-World orioles.
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Number of bird species: 3
(As at October 2025)
According to the recently (2025) amalgamated AviLidst, there are three species, in three genera in the Modulatricidae family. They are:
Spot-throat Modulatrix stictigula
Grey-chested Babbler Kakamega poliothorax
Dapple-throat Arcanator orostruthus
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Modulatricidae
Family AccountThese three brown, thrush-like birds are limited to small isolated areas of montane forest in tropical Africa. -
Modulatricidae
Family AccountModulatricidae is a small family of passerine birds which are restricted to Africa. These species have been taxonomic enigmas in the past.
Given that this is a tiny family with just three species albeit in three genera, Fatbirder provides active links below to all species.
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Dapple-throat Arcanator orostruthus
Species AccountSupreme understory skulker in montane forest, resembling a hybrid between a thrush and babbler. A dull and dowdy understory specialist with brown-olive... -
Dapple-throat Arcanator orostruthus
Species AccountThe dapple-throat or dappled mountain-robin (Arcanator orostruthus) is a species of bird in the Arcanatoridae family. It is found in Mozambique and Tanzania. Its natural habitat is subtropical or tropical moist montane forests. It is threatened by habitat loss. -
Dapple-throat Arcanator orostruthus
Species AccountArcanator orostruthus is listed as Near Threatened under criteria C2a(i). -
Dapple-throat Arcanator orostruthus
Species AccountSound archive and distribution map -
Grey-chested Kakamega Kakamega poliothorax
Species AccountUnusual thrush-like bird with bright reddish-brown upperparts and gray underparts. Found in the understory of montane forest, often in thick and moist areas -
Grey-chested Kakamega Kakamega poliothorax
Species AccountThe grey-chested babbler (Kakamega poliothorax), also known as the grey-chested illadopsis, is a species of bird in the Arcanatoridae family. It is the only species in its genus. It is found in Burundi, Cameroon, Democratic Republic of the Congo, Equatorial Guinea, Kenya, Nigeria, Rwanda, and Uganda. Its natural habitat is subtropical or tropical moist montane forests. -
Grey-chested Kakamega Kakamega poliothorax
Species AccountKakamega poliothorax is listed as Least Concern. -
Grey-chested Kakamega Kakamega poliothorax
Species AccountSound archive and distribution map -
Spot-throat Modulatrix stictigula
Species AccountSupreme understory skulker in montane forest, resembling a hybrid between a thrush and babbler. A dull but distinctive-looking bird if seen well. -
Spot-throat Modulatrix stictigula
Species AccountThe spot-throat (Modulatrix stictigula) is a species of bird in the Arcanatoridae family. It is found in northern Malawi and Tanzania. Its natural habitat is subtropical or tropical moist montane forests. -
Spot-throat Modulatrix stictigula
Species AccountSound archive and distribution map